Indonesian zoo staff have apologized after being condemned by animal rights activist when footage of an orangutan smoking went viral.

A visitor allegedly flicked a cigarette into the enclosure at Bandung zoo, and Odon, 22, picked it up and started puffing - adding to Indonesia's bad record of animal protection.

Marison Guciano, founder of the Indonesia Animal Welfare Society, attacked the zoo's management for "weak control", but also criticised a wider lack of welfare standards at zoos.